From ef2f6d90d20e8d261e88f7a78f204a185232619f Mon Sep 17 00:00:00 2001 From: depristo Date: Mon, 15 Nov 2010 22:19:22 +0000 Subject: [PATCH] VQSR now operates on LOD scores in the INFO field directly, and doesn't adjust the QUAL field. New format for tranches file uses LOD score. Old file format no longer supported. log10sumlog10() function, a very useful utility in MathUtils. No more ExtendedPileupElement! Robust math calculations in GMM so that no infinities are generated! HaplotypeScore refactored to enable use of filtered context. Not yet enabled... InferredContext getDouble and getInteger arguments now parse values from Strings if necessary git-svn-id: file:///humgen/gsa-scr1/gsa-engineering/svn_contents/trunk@4684 348d0f76-0448-11de-a6fe-93d51630548a --- .../annotator/AnnotatorInputTableCodec.java | 7 +- .../walkers/annotator/HaplotypeScore.java | 56 +- .../varianteval/VariantEvalWalker.java | 3 +- .../ApplyVariantCuts.java | 43 +- .../GenerateVariantClustersWalker.java | 3 +- .../walkers/variantrecalibration/Tranche.java | 49 +- .../variantrecalibration/VariantDatum.java | 4 +- .../VariantGaussianMixtureModel.java | 32 +- .../VariantRecalibrator.java | 36 +- .../genotyper/UnifiedGenotyperEngine.java | 6 +- .../broadinstitute/sting/utils/MathUtils.java | 13 +- .../pileup/AbstractReadBackedPileup.java | 13 - .../utils/pileup/ExtendedPileupElement.java | 29 - .../sting/utils/pileup/ReadBackedPileup.java | 3 - .../VariantEvalIntegrationTest.java | 4 +- .../VariantGaussianMixtureModelUnitTest.java | 40 +- ...ntRecalibrationWalkersIntegrationTest.java | 16 +- testdata/tranches.6.txt | 14 +- testdata/tranches.raw.dat | 36361 ++++++++-------- 19 files changed, 18347 insertions(+), 18385 deletions(-) delete mode 100755 java/src/org/broadinstitute/sting/utils/pileup/ExtendedPileupElement.java diff --git a/java/src/org/broadinstitute/sting/gatk/refdata/features/annotator/AnnotatorInputTableCodec.java b/java/src/org/broadinstitute/sting/gatk/refdata/features/annotator/AnnotatorInputTableCodec.java index 903133ffa..e87986a2e 100755 --- a/java/src/org/broadinstitute/sting/gatk/refdata/features/annotator/AnnotatorInputTableCodec.java +++ b/java/src/org/broadinstitute/sting/gatk/refdata/features/annotator/AnnotatorInputTableCodec.java @@ -92,13 +92,10 @@ public class AnnotatorInputTableCodec implements ReferenceDependentFeatureCodec< GenomeLoc loc; String chr = st.nextToken(); - if ( chr.indexOf(":") != -1 ) { + if ( chr.indexOf(":") != -1 ) loc = genomeLocParser.parseGenomeInterval(chr); - } else { - if ( st.countTokens() < 3 ) - throw new CodecLineParsingException("Couldn't parse GenomeLoc out of the following line because there aren't enough tokens.\nLine: " + line); + else loc = genomeLocParser.createGenomeLoc(chr, Integer.valueOf(st.nextToken()), Integer.valueOf(st.nextToken())); - } return new AnnotatorInputTableFeature(loc.getContig(), (int)loc.getStart(), (int)loc.getStop()); } diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/annotator/HaplotypeScore.java b/java/src/org/broadinstitute/sting/gatk/walkers/annotator/HaplotypeScore.java index b3288d738..d539007dd 100644 --- a/java/src/org/broadinstitute/sting/gatk/walkers/annotator/HaplotypeScore.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/annotator/HaplotypeScore.java @@ -40,6 +40,7 @@ import org.broadinstitute.sting.utils.pileup.*; import java.util.*; import net.sf.samtools.SAMRecord; import org.broadinstitute.sting.utils.sam.ReadUtils; +import org.broadinstitute.sting.utils.sam.GATKSAMRecord; public class HaplotypeScore implements InfoFieldAnnotation, StandardAnnotation { private final static boolean DEBUG = false; @@ -47,6 +48,11 @@ public class HaplotypeScore implements InfoFieldAnnotation, StandardAnnotation { private final static int MAX_CONSENSUS_HAPLOTYPES_TO_CONSIDER = 20; private final static char REGEXP_WILDCARD = '.'; + public boolean useRead(PileupElement p) { + return ! ReadUtils.is454Read(p.getRead()); + //return ! (p.getRead() instanceof GATKSAMRecord && ! ((GATKSAMRecord)p.getRead()).isGoodBase(p.getOffset())) && ! ReadUtils.is454Read(p.getRead()); + } + public Map annotate(RefMetaDataTracker tracker, ReferenceContext ref, Map stratifiedContexts, VariantContext vc) { if ( !vc.isBiallelic() || !vc.isSNP() || stratifiedContexts.size() == 0 ) // size 0 means that call was made by someone else and we have no data here return null; @@ -88,15 +94,11 @@ public class HaplotypeScore implements InfoFieldAnnotation, StandardAnnotation { ArrayList haplotypeList = new ArrayList(); PriorityQueue haplotypeQueue = new PriorityQueue(100, new HaplotypeComparator()); - - for ( ExtendedPileupElement p : pileup.extendedForeachIterator() ) { - if (ReadUtils.is454Read(p.getRead())) - continue; - Haplotype haplotypeFromRead = getHaplotypeFromRead(p, contextSize); - - - haplotypeQueue.add(haplotypeFromRead); - //haplotypeList.add(haplotypeFromRead); + for ( PileupElement p : pileup ) { + if ( useRead(p) ) { + Haplotype haplotypeFromRead = getHaplotypeFromRead(p, contextSize); + haplotypeQueue.add(haplotypeFromRead); + } } // Now that priority queue has been built with all reads at context, we need to merge and find possible segregating haplotypes @@ -161,7 +163,7 @@ public class HaplotypeScore implements InfoFieldAnnotation, StandardAnnotation { return null; } - private Haplotype getHaplotypeFromRead(ExtendedPileupElement p, int contextSize) { + private Haplotype getHaplotypeFromRead(PileupElement p, int contextSize) { SAMRecord read = p.getRead(); int readOffsetFromPileup = p.getOffset(); int baseOffsetStart = readOffsetFromPileup - (contextSize - 1)/2; @@ -239,24 +241,28 @@ public class HaplotypeScore implements InfoFieldAnnotation, StandardAnnotation { // if ( DEBUG ) System.out.printf("HAP1: %s%n", haplotypes.get(1)); double[][] haplotypeScores = new double[pileup.size()][haplotypes.size()]; - for ( ExtendedPileupElement p : pileup.extendedForeachIterator() ) { - SAMRecord read = p.getRead(); - int readOffsetFromPileup = p.getOffset(); + int pileupOffset = 0; + for ( PileupElement p : pileup ) { + if ( useRead(p) ) { + SAMRecord read = p.getRead(); + int readOffsetFromPileup = p.getOffset(); - if (ReadUtils.is454Read(read)) - continue; + if (ReadUtils.is454Read(read)) + continue; - if ( DEBUG ) System.out.printf("--------------------------------------------- Read %s%n", read.getReadName()); - double m = 10000000; - for ( int i = 0; i < haplotypes.size(); i++ ) { - Haplotype haplotype = haplotypes.get(i); - int start = readOffsetFromPileup - (contextSize - 1)/2; - double score = scoreReadAgainstHaplotype(read, start, contextSize, haplotype); - haplotypeScores[p.getPileupOffset()][i] = score; - if ( DEBUG ) System.out.printf(" vs. haplotype %d = %f%n", i, score); - m = Math.min(score, m); + if ( DEBUG ) System.out.printf("--------------------------------------------- Read %s%n", read.getReadName()); + double m = 10000000; + for ( int i = 0; i < haplotypes.size(); i++ ) { + Haplotype haplotype = haplotypes.get(i); + int start = readOffsetFromPileup - (contextSize - 1)/2; + double score = scoreReadAgainstHaplotype(read, start, contextSize, haplotype); + haplotypeScores[pileupOffset][i] = score; + if ( DEBUG ) System.out.printf(" vs. haplotype %d = %f%n", i, score); + m = Math.min(score, m); + } + if ( DEBUG ) System.out.printf(" => best score was %f%n", m); } - if ( DEBUG ) System.out.printf(" => best score was %f%n", m); + pileupOffset++; // todo -- remove me } double overallScore = 0.0; diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalWalker.java b/java/src/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalWalker.java index 7cfdbcafa..11398fb04 100755 --- a/java/src/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalWalker.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalWalker.java @@ -43,6 +43,7 @@ import org.broadinstitute.sting.gatk.walkers.RodWalker; import org.broadinstitute.sting.gatk.walkers.Window; import org.broadinstitute.sting.gatk.walkers.TreeReducible; import org.broadinstitute.sting.gatk.walkers.variantrecalibration.Tranche; +import org.broadinstitute.sting.gatk.walkers.variantrecalibration.VariantRecalibrator; import org.broadinstitute.sting.utils.SampleUtils; import org.broadinstitute.sting.utils.report.ReportMarshaller; import org.broadinstitute.sting.utils.report.VE2ReportFactory; @@ -307,7 +308,7 @@ public class VariantEvalWalker extends RodWalker implements Tr // we are going to build a few select names automatically from the tranches file for ( Tranche t : Tranche.readTraches(new File(TRANCHE_FILENAME)) ) { logger.info("Adding select for all variant above the pCut of : " + t); - SELECT_EXPS.add(String.format("QUAL >= %.2f", t.pCut)); + SELECT_EXPS.add(String.format(VariantRecalibrator.VQS_LOD_KEY + " >= %.2f", t.minVQSLod)); SELECT_NAMES.add(String.format("FDR-%.2f", t.fdr)); } } diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/ApplyVariantCuts.java b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/ApplyVariantCuts.java index d4e0153d6..1da542f5e 100755 --- a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/ApplyVariantCuts.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/ApplyVariantCuts.java @@ -100,11 +100,11 @@ public class ApplyVariantCuts extends RodWalker { if( tranches.size() >= 2 ) { for( int iii = 0; iii < tranches.size() - 1; iii++ ) { Tranche t = tranches.get(iii); - hInfo.add(new VCFFilterHeaderLine(t.name, String.format("FDR tranche level at qual: " + t.pCut + " <= x < " + tranches.get(iii+1).pCut))); + hInfo.add(new VCFFilterHeaderLine(t.name, String.format("FDR tranche level at VSQ Lod: " + t.minVQSLod + " <= x < " + tranches.get(iii+1).minVQSLod))); } } if( tranches.size() >= 1 ) { - hInfo.add(new VCFFilterHeaderLine(tranches.get(0).name + "+", String.format("FDR tranche level at qual < " + tranches.get(0).pCut))); + hInfo.add(new VCFFilterHeaderLine(tranches.get(0).name + "+", String.format("FDR tranche level at VQS Lod < " + tranches.get(0).minVQSLod))); } else { throw new UserException("No tranches were found in the file or were above the FDR Filter level " + FDR_FILTER_LEVEL); } @@ -131,28 +131,31 @@ public class ApplyVariantCuts extends RodWalker { if( vc != null && !vc.getSource().equals(DbSNPHelper.STANDARD_DBSNP_TRACK_NAME) && vc.isSNP() ) { String filterString = null; if( !vc.isFiltered() ) { - final double qual = vc.getPhredScaledQual(); - - for( int i = tranches.size() - 1; i >= 0; i-- ) { - Tranche tranche = tranches.get(i); - if( qual >= tranche.pCut ) { - if (i == tranches.size() - 1) { - filterString = VCFConstants.PASSES_FILTERS_v4; - } else { - filterString = tranche.name; + try { + final double lod = vc.getAttributeAsDouble(VariantRecalibrator.VQS_LOD_KEY); + for( int i = tranches.size() - 1; i >= 0; i-- ) { + Tranche tranche = tranches.get(i); + if( lod >= tranche.minVQSLod ) { + if (i == tranches.size() - 1) { + filterString = VCFConstants.PASSES_FILTERS_v4; + } else { + filterString = tranche.name; + } + break; } - break; } - } - if( filterString == null ) { - filterString = tranches.get(0).name+"+"; - } + if( filterString == null ) { + filterString = tranches.get(0).name+"+"; + } - if ( !filterString.equals(VCFConstants.PASSES_FILTERS_v4) ) { - Set filters = new HashSet(); - filters.add(filterString); - vc = VariantContext.modifyFilters(vc, filters); + if ( !filterString.equals(VCFConstants.PASSES_FILTERS_v4) ) { + Set filters = new HashSet(); + filters.add(filterString); + vc = VariantContext.modifyFilters(vc, filters); + } + } catch ( ClassCastException e ) { + throw new UserException.MalformedFile(vc.getSource(), "Invalid value for VQS key " + VariantRecalibrator.VQS_LOD_KEY + " at variant " + vc, e); } } diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/GenerateVariantClustersWalker.java b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/GenerateVariantClustersWalker.java index c82c18c09..b2907e3cf 100755 --- a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/GenerateVariantClustersWalker.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/GenerateVariantClustersWalker.java @@ -183,7 +183,6 @@ public class GenerateVariantClustersWalker extends RodWalker vcsHapMap = tracker.getVariantContexts(ref, "hapmap", null, context.getLocation(), false, true); @@ -201,7 +200,7 @@ public class GenerateVariantClustersWalker extends RodWalker 0.0 && variantDatum.qual > QUAL_THRESHOLD ) { + if( variantDatum.weight > 0.0 && vc.getPhredScaledQual() > QUAL_THRESHOLD ) { mapList.add( variantDatum ); } } diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/Tranche.java b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/Tranche.java index 1be2eb1d0..2caf27e8a 100755 --- a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/Tranche.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/Tranche.java @@ -49,33 +49,35 @@ import java.util.*; public class Tranche implements Comparable { private static final int CURRENT_VERSION = 2; - public double fdr, pCut, targetTiTv, knownTiTv, novelTiTv; + public double fdr, minVQSLod, targetTiTv, knownTiTv, novelTiTv; public int numKnown,numNovel; public String name; - public Tranche(double fdr, double targetTiTv, double pCut, int numKnown, double knownTiTv, int numNovel, double novelTiTv) { - this(fdr, targetTiTv, pCut, numKnown, knownTiTv, numNovel, novelTiTv, "anonymous"); + public Tranche(double fdr, double targetTiTv, double minVQSLod, int numKnown, double knownTiTv, int numNovel, double novelTiTv) { + this(fdr, targetTiTv, minVQSLod, numKnown, knownTiTv, numNovel, novelTiTv, "anonymous"); } - public Tranche(double fdr, double targetTiTv, double pCut, int numKnown, double knownTiTv, int numNovel, double novelTiTv, String name) { + public Tranche(double fdr, double targetTiTv, double minVQSLod, int numKnown, double knownTiTv, int numNovel, double novelTiTv, String name) { this.fdr = fdr; this.targetTiTv = targetTiTv; - this.pCut = pCut; + this.minVQSLod = minVQSLod; this.novelTiTv = novelTiTv; this.numNovel = numNovel; this.knownTiTv = knownTiTv; this.numKnown = numKnown; this.name = name; -// if ( targetTiTv < 0.5 || targetTiTv > 10 ) -// throw new UserException("Target Ti/Tv ratio is unreasonable " + targetTiTv); -// -// if ( numKnown < 0 || numNovel < 0) -// throw new ReviewedStingException("Invalid tranch - no. variants is < 0 : known " + numKnown + " novel " + numNovel); + if ( fdr <= 0.0 ) + throw new UserException("Target FDR is unreasonable " + fdr); + + if ( targetTiTv < 0.5 || targetTiTv > 10 ) + throw new UserException("Target Ti/Tv ratio is unreasonable " + targetTiTv); + + if ( numKnown < 0 || numNovel < 0) + throw new ReviewedStingException("Invalid tranch - no. variants is < 0 : known " + numKnown + " novel " + numNovel); if ( name == null ) throw new ReviewedStingException("BUG -- name cannot be null"); - } public int compareTo(Tranche other) { @@ -83,8 +85,8 @@ public class Tranche implements Comparable { } public String toString() { - return String.format("Tranche fdr=%.2f minQual=%.2f known=(%d @ %.2f) novel=(%d @ %.2f) name=%s]", - fdr, pCut, numKnown, knownTiTv, numNovel, novelTiTv, name); + return String.format("Tranche fdr=%.2f minVQSLod=%.4f known=(%d @ %.2f) novel=(%d @ %.2f) name=%s]", + fdr, minVQSLod, numKnown, knownTiTv, numNovel, novelTiTv, name); } /** @@ -103,12 +105,12 @@ public class Tranche implements Comparable { stream.println("# Variant quality score tranches file"); stream.println("# Version number " + CURRENT_VERSION); - stream.println("FDRtranche,targetTiTv,numKnown,numNovel,knownTiTv,novelTiTv,pCut,filterName"); + stream.println("FDRtranche,targetTiTv,numKnown,numNovel,knownTiTv,novelTiTv,minVQSLod,filterName"); Tranche prev = null; for ( Tranche t : tranches ) { - stream.printf("%.2f,%.2f,%d,%d,%.4f,%.4f,%.2f,FDRtranche%.2fto%.2f%n", - t.fdr,t.targetTiTv,t.numKnown,t.numNovel,t.knownTiTv,t.novelTiTv, t.pCut, + stream.printf("%.2f,%.2f,%d,%d,%.4f,%.4f,%.4f,FDRtranche%.2fto%.2f%n", + t.fdr,t.targetTiTv,t.numKnown,t.numNovel,t.knownTiTv,t.novelTiTv, t.minVQSLod, (prev == null ? 0.0 : prev.fdr), t.fdr); prev = t; } @@ -117,8 +119,10 @@ public class Tranche implements Comparable { } private static double getDouble(Map bindings, String key, boolean required) { - if ( bindings.containsKey(key) ) - return Double.valueOf(bindings.get(key)); + if ( bindings.containsKey(key) ) { + String val = bindings.get(key); + return Double.valueOf(val); + } else if ( required ) { throw new UserException.MalformedFile("Malformed tranches file. Missing required key " + key); } @@ -154,6 +158,11 @@ public class Tranche implements Comparable { final String[] vals = line.split(","); if( header == null ) { header = vals; + if ( header.length == 5 ) + // old style tranches file, throw an error + throw new UserException.MalformedFile(f, "Unfortuanately, your tranches file is from a previous version of this tool and cannot be used with the latest code. Please rerun VariantRecalibrator"); + if ( header.length != 8 ) + throw new UserException.MalformedFile(f, "Expected 8 elements in header line " + line); } else { if ( header.length != vals.length ) throw new UserException.MalformedFile(f, "Line had too few/many fields. Header = " + header.length + " vals " + vals.length + " line " + line); @@ -162,11 +171,11 @@ public class Tranche implements Comparable { for ( int i = 0; i < vals.length; i++ ) bindings.put(header[i], vals[i]); tranches.add(new Tranche(getDouble(bindings,"FDRtranche", true), getDouble(bindings,"targetTiTv", false), - getDouble(bindings,"pCut", true), + getDouble(bindings,"minVQSLod", true), getInteger(bindings,"numKnown", false), getDouble(bindings,"knownTiTv", false), getInteger(bindings,"numNovel", true), - Math.max(getDouble(bindings,"novelTiTv", false), getDouble(bindings,"novelTITV", false)), + getDouble(bindings,"novelTiTv", true), bindings.get("filterName"))); } } diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantDatum.java b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantDatum.java index dc2ba54da..f49cd4b7a 100755 --- a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantDatum.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantDatum.java @@ -36,10 +36,10 @@ public class VariantDatum implements Comparable { public double[] annotations; public boolean isTransition; public boolean isKnown; - public double qual; + public double lod; public double weight; public int compareTo(VariantDatum other) { - return Double.compare(this.qual, other.qual); + return Double.compare(this.lod, other.lod); } } diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModel.java b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModel.java index 6600470aa..3254e4184 100755 --- a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModel.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModel.java @@ -444,7 +444,7 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel } public final double evaluateVariant( GenomeLocParser genomeLocParser, final VariantContext vc ) { - final double[] pVarInCluster = new double[maxGaussians]; + final double[] log10pVarInCluster = new double[maxGaussians]; final double[] annotations = new double[dataManager.numAnnotations]; for( int jjj = 0; jjj < dataManager.numAnnotations; jjj++ ) { @@ -452,14 +452,7 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel annotations[jjj] = (value - dataManager.meanVector[jjj]) / dataManager.varianceVector[jjj]; } - evaluateGaussiansForSingleVariant( annotations, pVarInCluster ); - - double sum = 0.0; - for( int kkk = 0; kkk < maxGaussians; kkk++ ) { - sum += pVarInCluster[kkk]; - } - - return sum; + return evaluateGaussiansForSingleVariant( annotations, log10pVarInCluster ); } // --------------------------------------------------------------------------------------------------------- @@ -474,7 +467,7 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel public final static List findTranches( final VariantDatum[] data, final double[] FDRtranches, double targetTiTv, File debugFile ) { logger.info(String.format("Finding tranches for %d variants with %d FDRs and a target TiTv of %.2f", data.length, FDRtranches.length, targetTiTv)); - List tranchesData = sortVariantsbyQual(data); + List tranchesData = sortVariantsbyLod(data); double[] runningTiTv = calculateRunningTiTv(tranchesData); if ( debugFile != null) { writeTranchesDebuggingInfo(debugFile, tranchesData, runningTiTv); } @@ -502,14 +495,14 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel out.println("Qual isTransition runningTiTv"); for ( int i = 0; i < runningTiTv.length; i++ ) { VariantDatum d = tranchesData.get(i); - out.printf("%.4f %d %.4f%n", d.qual, d.isTransition ? 1 : 0, runningTiTv[i]); + out.printf("%.4f %d %.4f%n", d.lod, d.isTransition ? 1 : 0, runningTiTv[i]); } } catch (FileNotFoundException e) { throw new UserException.CouldNotCreateOutputFile(f, e); } } - private final static List sortVariantsbyQual(final VariantDatum[] data) { + private final static List sortVariantsbyLod(final VariantDatum[] data) { List sorted = new ArrayList(Arrays.asList(data)); Collections.sort(sorted); return sorted; @@ -555,10 +548,10 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel public final static Tranche trancheOfVariants( final List data, int minI, double fdr, double targetTiTv ) { int numKnown = 0, numNovel = 0, knownTi = 0, knownTv = 0, novelTi = 0, novelTv = 0; - double qualThreshold = data.get(minI).qual; + double minLod = data.get(minI).lod; VariantDatum last = null; for ( VariantDatum datum : data ) { - if ( datum.qual >= qualThreshold ) { + if ( datum.lod >= minLod ) { //if( ! datum.isKnown ) System.out.println(datum.pos); if ( datum.isKnown ) { numKnown++; @@ -575,7 +568,7 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel double knownTiTv = knownTi / Math.max(1.0 * knownTv, 1.0); double novelTiTv = novelTi / Math.max(1.0 * novelTv, 1.0); - return new Tranche(fdr, targetTiTv, qualThreshold, numKnown, knownTiTv, numNovel, novelTiTv); + return new Tranche(fdr, targetTiTv, minLod, numKnown, knownTiTv, numNovel, novelTiTv); } public final static double fdrToTiTv(double desiredFDR, double targetTiTv) { @@ -666,8 +659,7 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel return likelihood / sumWeight; } - - private void evaluateGaussiansForSingleVariant( final double[] annotations, final double[] pVarInCluster ) { + private double evaluateGaussiansForSingleVariant( final double[] annotations, final double[] log10pVarInCluster ) { final int numAnnotations = annotations.length; @@ -689,10 +681,12 @@ public final class VariantGaussianMixtureModel extends VariantOptimizationModel final double denomLog10 = CONSTANT_GAUSSIAN_DENOM_LOG10 + sqrtDeterminantLog10[kkk]; final double evalGaussianPDFLog10 = (( -0.5 * sum ) / Math.log(10.0)) - denomLog10; - final double pVar1 = Math.pow(10.0, pClusterLog10[kkk] + evalGaussianPDFLog10); - pVarInCluster[kkk] = pVar1; + //final double pVar1 = Math.pow(10.0, pClusterLog10[kkk] + evalGaussianPDFLog10); + final double pVar1 = pClusterLog10[kkk] + evalGaussianPDFLog10; + log10pVarInCluster[kkk] = pVar1; } + return MathUtils.log10sumLog10(log10pVarInCluster); } diff --git a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrator.java b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrator.java index 958b98be8..0f9125690 100755 --- a/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrator.java +++ b/java/src/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrator.java @@ -40,6 +40,7 @@ import org.broadinstitute.sting.utils.*; import org.broadinstitute.sting.utils.collections.ExpandingArrayList; import org.broadinstitute.sting.utils.collections.NestedHashMap; import org.broadinstitute.sting.utils.exceptions.UserException; +import org.broadinstitute.sting.utils.exceptions.ReviewedStingException; import org.broadinstitute.sting.utils.vcf.VCFUtils; import java.io.File; @@ -57,6 +58,8 @@ import java.util.*; */ public class VariantRecalibrator extends RodWalker, ExpandingArrayList> { + public static final String VQS_LOD_KEY = "VQSLOD"; + public static final double SMALLEST_LOG10_PVAR = -1e6; ///////////////////////////// // Inputs @@ -115,13 +118,9 @@ public class VariantRecalibrator extends RodWalker inputNames = new HashSet(); private NestedHashMap priorCache = new NestedHashMap(); private boolean trustACField = false; - private double maxQualObserved = 0.0; private PrintStream tranchesStream = null; - private static double round2(double num) { - double result = num * 100.0; + private static double round4(double num) { + double result = num * 10000.0; result = Math.round(result); - result = result / 100.0; + result = result / 10000.0; return result; } @@ -196,8 +194,7 @@ public class VariantRecalibrator extends RodWalker hInfo = new HashSet(); final TreeSet samples = new TreeSet(); hInfo.addAll(VCFUtils.getHeaderFields(getToolkit(), inputNames)); - hInfo.add(new VCFInfoHeaderLine("OQ", 1, VCFHeaderLineType.Float, "The original variant quality score")); - hInfo.add(new VCFInfoHeaderLine("LOD", 1, VCFHeaderLineType.Float, "The log odds ratio calculated by the VR algorithm which was turned into the phred scaled recalibrated quality score")); + hInfo.add(new VCFInfoHeaderLine(VQS_LOD_KEY, 1, VCFHeaderLineType.Float, "log10-scaled probability of variant being true under the trained gaussian mixture model")); samples.addAll(SampleUtils.getUniqueSamplesFromRods(getToolkit(), inputNames)); final VCFHeader vcfHeader = new VCFHeader(hInfo, samples); @@ -288,19 +285,20 @@ public class VariantRecalibrator extends RodWalker maxQualObserved ) { - maxQualObserved = variantDatum.qual; + final double log10pVar = theModel.evaluateVariant( ref.getGenomeLocParser(),vc ); + final double lod = priorLodFactor + log10pVar; + + if ( lod == Double.NEGATIVE_INFINITY ) { + throw new ReviewedStingException("Negative infinity detected during summation at " + vc); } + variantDatum.pos = vc.getStart(); + variantDatum.lod = round4(lod); + mapList.add( variantDatum ); final Map attrs = new HashMap(vc.getAttributes()); - attrs.put("OQ", String.format("%.2f", vc.getPhredScaledQual())); - attrs.put("LOD", String.format("%.4f", lod)); - VariantContext newVC = VariantContext.modifyPErrorFiltersAndAttributes(vc, variantDatum.qual / 10.0, new HashSet(), attrs); + attrs.put(VariantRecalibrator.VQS_LOD_KEY, String.format("%.4f", lod)); + VariantContext newVC = VariantContext.modifyPErrorFiltersAndAttributes(vc, vc.getNegLog10PError(), new HashSet(), attrs); vcfWriter.add( newVC, ref.getBase() ); } diff --git a/java/src/org/broadinstitute/sting/playground/gatk/walkers/genotyper/UnifiedGenotyperEngine.java b/java/src/org/broadinstitute/sting/playground/gatk/walkers/genotyper/UnifiedGenotyperEngine.java index 5d74d5201..a398bbc4a 100755 --- a/java/src/org/broadinstitute/sting/playground/gatk/walkers/genotyper/UnifiedGenotyperEngine.java +++ b/java/src/org/broadinstitute/sting/playground/gatk/walkers/genotyper/UnifiedGenotyperEngine.java @@ -233,7 +233,7 @@ public class UnifiedGenotyperEngine { // the overall lod //double overallLog10PofNull = log10AlleleFrequencyPosteriors.get()[0]; - double overallLog10PofF = MathUtils.log10sum(log10AlleleFrequencyPosteriors.get(), 1); + double overallLog10PofF = MathUtils.log10sumLog10(log10AlleleFrequencyPosteriors.get(), 1); if ( DEBUG_SLOD ) System.out.println("overallLog10PofF=" + overallLog10PofF); // the forward lod @@ -243,7 +243,7 @@ public class UnifiedGenotyperEngine { afcm.get().getLog10PNonRef(tracker, refContext, GLs, log10AlleleFrequencyPriors, log10AlleleFrequencyPosteriors.get()); //double[] normalizedLog10Posteriors = MathUtils.normalizeFromLog10(log10AlleleFrequencyPosteriors.get(), true); double forwardLog10PofNull = log10AlleleFrequencyPosteriors.get()[0]; - double forwardLog10PofF = MathUtils.log10sum(log10AlleleFrequencyPosteriors.get(), 1); + double forwardLog10PofF = MathUtils.log10sumLog10(log10AlleleFrequencyPosteriors.get(), 1); if ( DEBUG_SLOD ) System.out.println("forwardLog10PofNull=" + forwardLog10PofNull + ", forwardLog10PofF=" + forwardLog10PofF); // the reverse lod @@ -253,7 +253,7 @@ public class UnifiedGenotyperEngine { afcm.get().getLog10PNonRef(tracker, refContext, GLs, log10AlleleFrequencyPriors, log10AlleleFrequencyPosteriors.get()); //normalizedLog10Posteriors = MathUtils.normalizeFromLog10(log10AlleleFrequencyPosteriors.get(), true); double reverseLog10PofNull = log10AlleleFrequencyPosteriors.get()[0]; - double reverseLog10PofF = MathUtils.log10sum(log10AlleleFrequencyPosteriors.get(), 1); + double reverseLog10PofF = MathUtils.log10sumLog10(log10AlleleFrequencyPosteriors.get(), 1); if ( DEBUG_SLOD ) System.out.println("reverseLog10PofNull=" + reverseLog10PofNull + ", reverseLog10PofF=" + reverseLog10PofF); double forwardLod = forwardLog10PofF + reverseLog10PofNull - overallLog10PofF; diff --git a/java/src/org/broadinstitute/sting/utils/MathUtils.java b/java/src/org/broadinstitute/sting/utils/MathUtils.java index 77efeecc3..f7d7a434a 100755 --- a/java/src/org/broadinstitute/sting/utils/MathUtils.java +++ b/java/src/org/broadinstitute/sting/utils/MathUtils.java @@ -58,7 +58,7 @@ public class MathUtils { return s; } - public static double log10sum(double[] log10p, int start) { + public static double log10sumLog10(double[] log10p, int start) { double sum = 0.0; double maxValue = Utils.findMaxEntry(log10p); @@ -76,9 +76,14 @@ public class MathUtils { } public static double sumLog10(double[] log10values) { - double s = 0.0; - for ( double v : log10values) s += Math.pow(10.0, v); - return s; + return Math.pow(10.0, log10sumLog10(log10values)); +// double s = 0.0; +// for ( double v : log10values) s += Math.pow(10.0, v); +// return s; + } + + public static double log10sumLog10(double[] log10values) { + return log10sumLog10(log10values, 0); } public static boolean wellFormedDouble(double val) { diff --git a/java/src/org/broadinstitute/sting/utils/pileup/AbstractReadBackedPileup.java b/java/src/org/broadinstitute/sting/utils/pileup/AbstractReadBackedPileup.java index 95f1e3073..6ee64cabb 100644 --- a/java/src/org/broadinstitute/sting/utils/pileup/AbstractReadBackedPileup.java +++ b/java/src/org/broadinstitute/sting/utils/pileup/AbstractReadBackedPileup.java @@ -652,19 +652,6 @@ public abstract class AbstractReadBackedPileup extendedForeachIterator() { - ArrayList x = new ArrayList(size()); - int i = 0; - for ( PileupElement pile : this ) { - x.add(new ExtendedPileupElement(pile.getRead(), pile.getOffset(), i++, this)); - } - - return new IterableIterator(x.iterator()); - } - /** * Simple useful routine to count the number of deletion bases in this pileup * diff --git a/java/src/org/broadinstitute/sting/utils/pileup/ExtendedPileupElement.java b/java/src/org/broadinstitute/sting/utils/pileup/ExtendedPileupElement.java deleted file mode 100755 index a2ada94c5..000000000 --- a/java/src/org/broadinstitute/sting/utils/pileup/ExtendedPileupElement.java +++ /dev/null @@ -1,29 +0,0 @@ -package org.broadinstitute.sting.utils.pileup; - -import net.sf.samtools.SAMRecord; - -/** - * Created by IntelliJ IDEA. - * User: depristo - * Date: Apr 14, 2009 - * Time: 8:54:05 AM - * To change this template use File | Settings | File Templates. - */ -public class ExtendedPileupElement extends PileupElement { - private int pileupOffset = 0; - private ReadBackedPileup pileup = null; - - public ExtendedPileupElement( SAMRecord read, int readOffset, int pileupOffset, ReadBackedPileup pileup ) { - super(read, readOffset); - this.pileupOffset = pileupOffset; - this.pileup = pileup; - } - - public int getPileupOffset() { - return pileupOffset; - } - - public ReadBackedPileup getPileup() { - return pileup; - } -} \ No newline at end of file diff --git a/java/src/org/broadinstitute/sting/utils/pileup/ReadBackedPileup.java b/java/src/org/broadinstitute/sting/utils/pileup/ReadBackedPileup.java index 9454adfd7..ff6b2c122 100644 --- a/java/src/org/broadinstitute/sting/utils/pileup/ReadBackedPileup.java +++ b/java/src/org/broadinstitute/sting/utils/pileup/ReadBackedPileup.java @@ -141,9 +141,6 @@ public interface ReadBackedPileup extends Iterable { */ public ReadBackedPileup getPileupForSample(String sampleName); - // todo -- delete or make private - public IterableIterator extendedForeachIterator(); - /** * Simple useful routine to count the number of deletion bases in this pileup * diff --git a/java/test/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalIntegrationTest.java b/java/test/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalIntegrationTest.java index 9f84a3e65..912907721 100755 --- a/java/test/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalIntegrationTest.java +++ b/java/test/org/broadinstitute/sting/gatk/walkers/varianteval/VariantEvalIntegrationTest.java @@ -143,8 +143,8 @@ public class VariantEvalIntegrationTest extends WalkerTest { @Test public void testTranches() { - String extraArgs = "-T VariantEval -R "+ hg18Reference +" -B:eval,vcf " + validationDataLocation + "GA2.WEx.cleaned.ug.snpfiltered.indelfiltered.optimized.vcf -o %s -E TiTvVariantEvaluator -L chr1 -noStandard -reportType CSV -tf " + testDir + "tranches.4.txt"; - WalkerTestSpec spec = new WalkerTestSpec(extraArgs,1,Arrays.asList("85b6621c64cc8f9a3b68cea644edf216")); + String extraArgs = "-T VariantEval -R "+ hg18Reference +" -B:eval,vcf " + validationDataLocation + "GA2.WEx.cleaned.ug.snpfiltered.indelfiltered.optimized.vcf -o %s -E TiTvVariantEvaluator -L chr1 -noStandard -reportType CSV -tf " + testDir + "tranches.6.txt"; + WalkerTestSpec spec = new WalkerTestSpec(extraArgs,1,Arrays.asList("1002c7a71e6c14f87398d924ac1d92af")); executeTestParallel("testTranches",spec); //executeTest("testACDiscordanceAtAC1EvalAC2Comp",spec); } diff --git a/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModelUnitTest.java b/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModelUnitTest.java index b0933735c..50d1484f9 100644 --- a/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModelUnitTest.java +++ b/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantGaussianMixtureModelUnitTest.java @@ -59,7 +59,7 @@ public final class VariantGaussianMixtureModelUnitTest extends BaseTest { VariantDatum[] variantData1 = new VariantDatum[N_VARIANTS]; private final File QUAL_DATA = new File(testDir + "tranches.raw.dat"); - private final double[] FDRS = new double[]{0.1, 1, 10, 100}; + private final double[] FDRS = new double[]{0.1, 1, 2, 10}; private final double TARGET_TITV = 2.8; private final File EXPECTED_TRANCHES_NEW = new File(testDir + "tranches.6.txt"); private final File EXPECTED_TRANCHES_OLD = new File(testDir + "tranches.4.txt"); @@ -68,12 +68,13 @@ public final class VariantGaussianMixtureModelUnitTest extends BaseTest { List vd = new ArrayList(); try { for ( String line : new XReadLines(QUAL_DATA, true) ) { - String[] parts = line.split(" "); + String[] parts = line.split("\t"); + // QUAL,TRANSITION,ID,LOD,FILTER if ( ! parts[0].equals("QUAL") ) { VariantDatum datum = new VariantDatum(); - datum.qual = Double.valueOf(parts[0]); + datum.lod = Double.valueOf(parts[3]); datum.isTransition = parts[1].equals("1"); - datum.isKnown = parts[2].equals("1"); + datum.isKnown = ! parts[2].equals("."); vd.add(datum); } } @@ -94,7 +95,7 @@ public final class VariantGaussianMixtureModelUnitTest extends BaseTest { read(EXPECTED_TRANCHES_NEW); } - @Test + @Test(expectedExceptions = {UserException.MalformedFile.class}) public final void readOldFormat() { read(EXPECTED_TRANCHES_OLD); } @@ -103,13 +104,6 @@ public final class VariantGaussianMixtureModelUnitTest extends BaseTest { return Tranche.readTraches(f); } - @Test - public final void testNewAndOldAretheSame() { - List newFormat = read(EXPECTED_TRANCHES_NEW); - List oldFormat = read(EXPECTED_TRANCHES_OLD); - assertTranchesAreTheSame(newFormat, oldFormat, false, true); - } - private static void assertTranchesAreTheSame(List newFormat, List oldFormat, boolean completeP, boolean includeName) { Assert.assertEquals(oldFormat.size(), newFormat.size()); for ( int i = 0; i < newFormat.size(); i++ ) { @@ -136,15 +130,15 @@ public final class VariantGaussianMixtureModelUnitTest extends BaseTest { assertTranchesAreTheSame(read(EXPECTED_TRANCHES_NEW), tranches, true, false); } -// @Test(expectedExceptions = {UserException.class}) -// public final void testBadFDR() { -// List vd = readData(); -// VariantGaussianMixtureModel.findTranches(vd.toArray(new VariantDatum[0]), new double[]{-1}, TARGET_TITV); -// } -// -// @Test(expectedExceptions = {UserException.class}) -// public final void testBadTargetTiTv() { -// List vd = readData(); -// VariantGaussianMixtureModel.findTranches(vd.toArray(new VariantDatum[0]), FDRS, 0.1); -// } + @Test(expectedExceptions = {UserException.class}) + public final void testBadFDR() { + List vd = readData(); + VariantGaussianMixtureModel.findTranches(vd.toArray(new VariantDatum[0]), new double[]{-1}, TARGET_TITV); + } + + @Test(expectedExceptions = {UserException.class}) + public final void testBadTargetTiTv() { + List vd = readData(); + VariantGaussianMixtureModel.findTranches(vd.toArray(new VariantDatum[0]), FDRS, 0.1); + } } diff --git a/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrationWalkersIntegrationTest.java b/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrationWalkersIntegrationTest.java index 30d44a77e..0024149bb 100755 --- a/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrationWalkersIntegrationTest.java +++ b/java/test/org/broadinstitute/sting/gatk/walkers/variantrecalibration/VariantRecalibrationWalkersIntegrationTest.java @@ -30,15 +30,15 @@ public class VariantRecalibrationWalkersIntegrationTest extends WalkerTest { VRTest yriTrio = new VRTest("yri.trio.gatk_glftrio.intersection.annotated.filtered.chr1.vcf", "ab2629d67e378fd3aceb8318f0fbfe04", // in vcf - "4dd95e9d8e5d21a6ab73de67d9663492", // tranches - "4e893672230fca625f70b0491f3b36cb", // recalVCF - "371b0e2796982485ae050e46892f6660"); // cut VCF + "d8f0a92aa42989d332d3c65f61352978", // tranches + "3370e51f6e5035f2472c66762d13c821", // recalVCF + "4801e6cfaeadc81c5c1f9bf5948e2a1c"); // cut VCF VRTest lowPass = new VRTest("lowpass.N3.chr1.raw.vcf", "725489156426e4ddd8d623ab3d4b1023", // in vcf - "dfc07132f592a811d0d6c25a4cb67a09", // tranches - "d52e4f511c9c00f8c21dffea81c47103", // recalVCF - "39716b2f03b50e88855d7975dd1b9b3e"); // cut VCF + "3a7067247146f4a77bb4fd7bc36f94c4", // tranches + "9a525c6838bff695321fc7ac0e458f9c", // recalVCF + "41e0a16af150244454ee68948ced00fb"); // cut VCF @DataProvider(name = "VRTest") public Object[][] createData1() { @@ -64,7 +64,7 @@ public class VariantRecalibrationWalkersIntegrationTest extends WalkerTest { executeTest("testGenerateVariantClusters-"+params.inVCF, spec).getFirst(); } - @Test(dataProvider = "VRTest", dependsOnMethods = {"testGenerateVariantClusters"}, enabled = true) + @Test(dataProvider = "VRTest", enabled = true) public void testVariantRecalibrator(VRTest params) { //System.out.printf("PARAMS FOR %s is %s%n", vcf, clusterFile); WalkerTest.WalkerTestSpec spec = new WalkerTest.WalkerTestSpec( @@ -85,7 +85,7 @@ public class VariantRecalibrationWalkersIntegrationTest extends WalkerTest { executeTest("testVariantRecalibrator-"+params.inVCF, spec).getFirst(); } - @Test(dataProvider = "VRTest", dependsOnMethods = {"testVariantRecalibrator"}, enabled = true) + @Test(dataProvider = "VRTest", enabled = true) public void testApplyVariantCuts(VRTest params) { WalkerTest.WalkerTestSpec spec = new WalkerTest.WalkerTestSpec( "-R " + b36KGReference + diff --git a/testdata/tranches.6.txt b/testdata/tranches.6.txt index 3849718f4..983c444b4 100644 --- a/testdata/tranches.6.txt +++ b/testdata/tranches.6.txt @@ -1,7 +1,7 @@ -# comment 1 -# comment 2 -FDRtranche,targetTiTv,numKnown,numNovel,knownTiTv,novelTiTv,pCut,filterName -0.10,2.8000,15791,893,3.3086,2.8000,1.1800,FDRtranche0.00to0.10 -1.00,2.8000,15824,903,3.3047,2.7782,1.0600,FDRtranche0.10to1.00 -10.00,2.8000,16037,984,3.2937,2.5269,0.3800,FDRtranche1.00to10.00 -100.00,2.8000,16578,1602,3.2291,1.6091,0.0000,FDRtranche10.00to100.00 +# Variant quality score tranches file +# Version number 2 +FDRtranche,targetTiTv,numKnown,numNovel,knownTiTv,novelTiTv,minVQSLod,filterName +0.10,2.80,15839,897,3.3052,2.8008,-3.8485,FDRtranche0.00to0.10 +1.00,2.80,15844,900,3.3043,2.7815,-3.8652,FDRtranche0.10to1.00 +2.00,2.80,15847,902,3.3051,2.7583,-3.8826,FDRtranche1.00to2.00 +10.00,2.80,16153,1039,3.2744,2.5704,-5.1058,FDRtranche2.00to10.00 diff --git a/testdata/tranches.raw.dat b/testdata/tranches.raw.dat index f03c9142f..7090c21b1 100644 --- a/testdata/tranches.raw.dat +++ b/testdata/tranches.raw.dat @@ -1,18180 +1,18181 @@ -27.18 1 1 -38.05 1 1 -95.21 0 0 -2318.43 1 1 -3291.07 1 1 -10982.18 1 1 -93.97999999999999 0 1 -136.49 1 1 -357.08 1 1 -1040.28 1 1 -10.36 1 1 -144.53 1 1 -174.42000000000002 1 1 -6277.560000000001 1 1 -3.84 1 1 -282.35 1 1 -10.969999999999999 0 1 -28.0 1 1 -25.77 0 0 -0.0 1 0 -144.87 1 1 -481.99 1 1 -408.35 0 1 -135.64 1 1 -6985.76 1 1 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